Williams-Guillén, K., C. McCann, J.C. Martínez Sánchez, and F. Koontz. 2006. Resource availability and habitat use by mantled howling monkeys in a Nicaraguan coffee plantation: can agroforests serve as core habitat for a forest mammal? Animal Conservation 9: 331-338.
The incredible contact roars of Mantled Howler Monkeys (Alouatta palliata) are a frequent wake-up call to visitors to the New World tropics. They were studied in a Nicaraguan shade coffee plantation, which had a diverse (60+ species) canopy which insured that preferred foods were available to the monkeys year-round. The monkeys did not avoid areas of coffee plantations, but stuck to the larger trees. The authors concluded that shade coffee can serve as alternate wildlife habitat and corridors between forest fragments for howling monkeys and possibly other forest mammals.
